A conclusion on a nonuniform and discrete distribution of phene frequencies of occurrence is made on the basis of studies made for 39 characters of coloration (colour and pattern) in 13 668 Lacerta agilis L. samples from 135 points of the area of distribution. Numerous zones of a sharp difference in the frequencies of occurrence are found for certain phenes and their groups. Powerful phenetic boundaries formed by a sharp difference of frequencies simultaneously for many phenes are revealed and it is determined that these boundaries in many cases coincide fairly well with orographic regions. Different phenes are shown to mark the lizard natural groupings of various size: from separate populations to large geographical regions.